- Restricted growth, short internodes and small leaves. Initially, interveinal chlorotic blotches appear on mature leaves, but later symptoms spread upwards on the plant.
- The leaves eventually turn dull green or bronze, their edges turn down, and the plant remains dwarfed. Furthermore, bud and flower development at the top of the plant decreases.
- Copper deficiency can dramatically reduce yield (by 20-90%).
- The few fruits that are produced develop poorly with small, sunkenbrown areas scattered over their yellow-green skin.